1. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences , Beijing 100093 , China
2. China National Botanical Garden , Beijing 100093 , China
3. University of Chinese Academy of Sciences , Beijing 100049 , China
4. Missouri Botanical Garden , 4344 Shaw Boulevard, St. Louis, MO 63110 , USA
5. Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, Université des Antilles, EPHE , 57 rue Cuvier, CP39, Paris 75005 , France
6. Central Siberian Botanical Garden of the Siberian Branch of Russian Academy of Sciences , Zolotodolinskaya str. 101, Novosibirsk 630090 , Russia