1. State Key Laboratory of Herbage Improvement and Grassland Agro‐ecosystems, College of Ecology Lanzhou University Lanzhou China
2. State Key Laboratory of Plant Diversity and Specialty Crops, Institute of Botany Chinese Academy of Sciences Beijing China
3. China National Botanical Garden Beijing China
4. University of Chinese Academy of Sciences Beijing China
5. Missouri Botanical Garden St. Louis Missouri USA
6. Central Siberian Botanical Garden Russian Academy of Sciences Novosibirsk Russia
7. Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d'Histoire naturelle, CNRS Sorbonne Université, Université des Antilles Paris France