1. Korea Polar Research Institute (KOPRI), Incheon 21990, South Korea
2. Institut National de la Recherche Scientifique, Centre INRS-Institut Armand-Frappier, 531 boulevard de Prairies, Laval, Quebec, H7V 1B7, Canada
3. Department of Biology, Faculty of Science, Shinshu University, Matsumoto, Japan
4. Department of Biological Sciences, College of Natural Sciences, Seoul National University, Seoul 151–742, South Korea
5. Department of Life Science, Kyonggi University, Suwon 443–760, South Korea
6. Division of Agrifood and Environment, School of Water, Energy and Environment, Cranfield University, College Road, Cranfield MK43 0AL, UK