1. Department of Natural Sciences, Faculty of Science and Technology, Middlesex University , London NW4 4BT, UK
2. Institut Systématique Evolution Biodiversité (ISYEB), Sorbonne Université, MNHN, CNRS, EPHE , UA, 75005 Paris, France
3. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung , 10115 Berlin, Germany
4. German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, 04103 Leipzig , Germany
5. Institut Pasteur, Université Paris Cité, Sequence Bioinformatics , 75015 Paris, France