1. School of Medicine, University of St Andrews, North Haugh, St Andrews KY16 9TF, UK
2. Department of Plant Breeding, Swedish University of Agricultural Sciences, Lomma, SE-234 22, Sweden
3. Cell and Molecular Sciences, James Hutton Institute, Invergowrie, Dundee DD2 5DA, UK
4. Independent, Dundee DD5 4QT, UK