1. BGI-Shenzhen, Shenzhen, China
2. China National GeneBank, BGI-Shenzhen, Shenzhen, China
3. State Key Laboratory of Agricultural Genomics, BGI-Shenzhen, Shenzhen, China
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, China
5. Fairylake Botanical Garden, Shenzhen & Chinese Academy of Sciences, Shenzhen, China
6. School of Basic Medical, Qingdao University, China
7. BGI-Qingdao, BGI-Shenzhen, Qingdao, China
8. Hainan Key Laboratory for Sustainable Utilization of Tropical Bioresources, Institute of Tropical Agriculture and Forestry, Hainan University, Haikou, China
9. College of Chinese Medicine Materials, Jilin Agricultural University, China
10. MGI, BGI-Shenzhen, Shenzhen, China
11. Institute for Evolution and Biodiversity, University of Muenster, Germany†These authors contributed equally to this work