1. Global Institute for Food Security (GIFS), University of Saskatchewan , 421 Downey Rd, Saskatoon, SK S7N 4L8 , Canada
2. Department of Biology, University of Saskatchewan, 112 Science Place , Saskatoon, SK S7N 5E2 , Canada
3. Department of Plant Biology, College of Biological Sciences, University of California , Davis, CA 95616 , USA
4. Key Laboratory of Plant Resources, Institute of Botany, Chinese Academy of Sciences , Beijing 100093 , China