1. Global Institute for Food Security (GIFS), University of Saskatchewan, 421 Downey Rd, Saskatoon, SK S7N 4L8, Canada
2. Department of Biology, University of Saskatchewan, 112 Science Place, Saskatoon, SK S7N 5E2, Canada
3. Department of Plant Science, University of Saskatchewan, 51 Campus Drive, Saskatoon, SK S7N 5A8, Canada