1. Hawkesbury Institute for Environment, Western Sydney University, Hawkesbury Campus, Locked Bag 1797, Penrith, NSW 2751, Australia
2. Department of Biological and Environmental Sciences, University of Gothenburg, PO Box 461, SE-40530 Gothenburg, Sweden
3. ARC Centre of Excellence in Plant Energy Biology, Division of Plant Sciences, Research School of Biology, Building 134, The Australian National University, Canberra, ACT 2601, Australia