1. Department of Plant & Environmental New Resources, Kyung Hee University , Yongin 17104 , Republic of Korea
2. Forest Bioresources Department, National Institute of Forest Science , Suwon 16631 , Republic of Korea
3. Department of Horticulture, Michigan State University , East Lansing, MI 48824 , USA
4. DOE Great Lakes Bioenergy Research Center, Michigan State University , East Lansing, MI 48824 , USA
5. Department of Forestry, Michigan State University , East Lansing, MI 48824 , USA