1. State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University , Shanghai , China
2. School of Physical Sciences, University of Chinese Academy of Sciences , Beijing 100049 , China
3. Beijing Institutes of Life Science, Chinese Academy of Sciences , Beijing 100101 , China
4. University of Chinese Academy of Sciences , Beijing , China
5. Center for Excellence in Animal Evolution and Genetics, Chinese Academy of Sciences , Kunming , China