1. School of Biosciences , Cardiff CF10 3AX , UK
2. Cardiff University Neurosciences and Mental Health Research Institute, , Cardiff CF10 3AT , UK
3. Universitat Internacional de Catalunya Faculty of Medicine and Health Sciences Department of Basic Sciences, , Barcelona 08195 , Spain
4. Institut de Neurociènces, Universitat Autònoma de Barcelona , Bellaterra 08193 , Spain