1. School of Biological Sciences, University of Aberdeen , Aberdeen , UK
2. Shenzhen Key Laboratory of Metabolic Health, Center for Energy Metabolism and Reproduction, Shenzhen Institutes of Advanced Technology, Chinese Academy of Sciences , Shenzhen , P.R. China
3. State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences , Beijing , P.R. China
4. China Medical University , Shenyang, Liaoning , P.R. China