1. School of Life Sciences, University of Sussex, Falmer, Brighton BN1 9QG, UK
2. School of Plant Biology, The University of Western Australia, 35 Stirling Highway, Crawley, WA, Australia
3. Environmental Research Laboratory of the University of Arizona, 1601 East, Airport Drive, Tucson, AZ, USA
4. Faculty of Life Sciences and Computing, London Metropolitan University, London N7, UK
5. Department of Plant Sciences, College of Agricultural and Environmental Sciences, University of California, Davis, CA, USA