1. Kansas Cooperative Fish and Wildlife Research Unit, Division of Biology, Kansas State University, Manhattan, Kansas, USA
2. U.S. Geological Survey, Kansas Cooperative Fish and Wildlife Research Unit, Division of Biology, Kansas State University, Manhattan, Kansas, USA
3. Jonah Ventures, LLC, Manhattan, Kansas, USA
4. Colorado Parks and Wildlife Department, Lamar, Colorado, USA
5. Division of Biology, Kansas State University, Manhattan, Kansas, USA
6. Cooperative Institute for Research in Environmental Science, University of Colorado, Boulder, Colorado, USA