1. United Graduate School of Agricultural Sciences, Ehime University, Tarumi, Matsuyama, 790-8566, Japan
2. Graduate School of Bioagricultural Sciences, Nagoya University, Furo-cho Chikusa-ku, Nagoya, 464-8601, Japan
3. Department of Forest Resources, Faculty of Agriculture, Ehime University, Tarumi, Matsuyama 790-8566, Japan