1. University of Pittsburgh, Department of Biological Sciences, 4249 Fifth Avenue, Pittsburgh, PA 15260
2. 5 Smithsonian Tropical Research Institute, Luis Clement Avenue, Building 401 Tupper, Balboa Ancon, Postal 0843-03092, Panama; Fundación Yaguará Panamá, Edificio 181, Ciudad del Saber, Clayton, Ciudad de Panamá, Panamá