1. Department of Biological Sciences, Clemson University, Clemson, SC, United States of America
2. Plant and Environmental Sciences Department, Clemson University, Clemson, SC, United States of America
3. Smithsonian Marine Station at Fort Pierce, Smithsonian Institution, Fort Pierce, FL, United States of America
4. Departamento de Biologia Marina, Universidad Catolica del Norte, Coquimbo, IV Region, Chile