1. Australian Institute of Marine Science, Townsville, QLD, Australia
2. ARC Centre of Excellence for Mathematical & Statistical Frontiers, Queensland University of Technology, Brisbane, QLD, Australia
3. College of Science and Engineering, James Cook University, Townsville, QLD, Australia
4. School of Mathematical Sciences, Queensland University of Technology, Brisbane, QLD, Australia
5. ARC Centre of Excellence for Coral Reef Studies, College of Science and Engineering, James Cook University, Townsville, QLD, Australia