1. Tokyo University of Agriculture, Sakuragaoka 1-1-1, Setagaya, Tokyo 156-8502, Japan
2. GEVES, Station Nationale dâ–™Essais de Semences (SNES), Angers, France
3. Institute of Biological and Environmental Sciences, University of Aberdeen, AB24 3UU, UK
4. GEVES, Station Nationale d'Essais de Semences (SNES), Angers, France