1. Department of Sport and Health Sciences, College of Life and Environmental Sciences, University of Exeter, Exeter EX1 2LU, UK
2. Living Systems Institute, University of Exeter, Exeter EX4 4QD, UK
3. MRC-ARUK Centre of Research Excellence and National Institute of Health Research, Biomedical Research Centre, Postgraduate Entry Medical School, Royal Derby Hospital Centre, School of Medicine, University of Nottingham, Derby DE22 3DT, UK
4. Genetics of Complex Traits, University of Exeter Medical School, University of Exeter, Exeter EX1 2LU, UK
5. Department of Surgery, Postgraduate Entry Medical School, Royal Derby Hospital Centre, School of Medicine, University of Nottingham, Derby DE22 3DT, UK
6. Faculty of Health Sciences and Sport, University of Stirling, Stirling FK9 4LA, UK
7. School of Health Sciences, Örebro University, Örebro 70182, Sweden
8. Joint senior authorship