1. MRC-Versus Arthritis Centre for Musculoskeletal Ageing Research, University of Nottingham, Nottingham, UK
2. National Institute for Health Research (NIHR) Nottingham Biomedical Research Centre, Nottingham, UK
3. School of Medicine, University of Nottingham, Royal Derby Hospital Centre, Derby, UK
4. School of Biosciences and Phenome Centre Birmingham, University of Birmingham, Birmingham, Birmingham, UK
5. Beatson Institute for Cancer Research, Glasgow, UK
6. School of Life Sciences, University of Nottingham, Queens Medical Centre, Nottingham, UK
7. University of Stirling, Faculty of Health Sciences and Sport, Stirling, UK