1. Laboratory of Vaccine Materials Center for Vaccine and Adjuvant Research Laboratory of Gut Environmental System National Institutes of Biomedical Innovation Health and Nutrition (NIBIOHN) Osaka Japan
2. Graduate School of Pharmaceutical Sciences Osaka University Osaka Japan
3. Division of Applied Life Sciences Graduate School of Agriculture Kyoto University Kyoto Japan
4. Department of Dermatology Graduate School of Medicine Kyoto University Kyoto Japan
5. Graduate School of Medicine Kobe University Hyogo Japan
6. International Research and Development Center for Mucosal Vaccines The Institute of Medical Science The University of Tokyo Tokyo Japan
7. Graduate School of Medicine Graduate School of Dentistry Osaka University Osaka Japan