1. Institute of Medical Biology and Agency for Science Technology and Research (A*STAR), Singapore, Singapore.
2. Lee Kong Chian School of Medicine, Nanyang Technological University–Imperial College London, Singapore, Singapore.
3. Institute of Molecular and Cell Biology, Agency for Science Technology and Research (A*STAR), Singapore, Singapore.
4. Department of Orthopaedic Surgery, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.