1. Schepens Eye Research Institute of Mass Eye & Ear, Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, USA.
2. Paul F. Glenn Center for Biology of Aging Research, Department of Genetics, Blavatnik Institute, Harvard Medical School, Boston, Massachusetts, USA.
3. Whitehead Institute for Biomedical Research, Department of Biology, MIT, Cambridge, Massachusetts, USA.
4. Life Biosciences LLC, Boston, Massachusetts, USA.