1. Laboratory of Animal Reproduction, Department of Animal Science, Tokyo University of Agriculture, Atsugi, Japan.
2. NODAI Genome Research Center, Tokyo University of Agriculture, Tokyo, Japan.
3. Laboratory of Reproductive Physiology, Graduate School of Environmental and Life Science, Okayama University, Okayama, Japan.
4. Animal Feeding and Management Research Division, National Institute of Livestock and Grassland Science, Nasushiobara, Japan.