1. Department of Biology, University of Florida, Gainesville, Florida 32611‐8525 USA
2. Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611‐7800 USA
3. Department of Biology, Oberlin College, Oberlin, Ohio 44074‐1097 USA
4. Genetics Institute, University of Florida, Gainesville, Florida 32611 USA
5. Key Laboratory of Conservation Biology for Endangered Wildlife of the Ministry of Education, and Laboratory of Systematic and Evolutionary Botany and Biodiversity, College of Life Sciences, Zhejiang University, Hangzhou 310058 People's Republic of China
6. Department of Plant Sciences, University of Cambridge, Cambridge CB13EA United Kingdom