1. Department of Integrative Biology, University of California, Berkeley, 3040 Valley Life Sciences Building #3140, Berkeley, California 94720 USA
2. University and Jepson Herbaria, University of California, Berkeley, 1001 Valley Life Sciences Building #2465, Berkeley, California 94720 USA
3. Department of Plant and Microbial Biology, University of California, Berkeley, 111 Koshland Hall, Berkeley, California 94720 USA
4. School of Integrative Plant Sciences, Section of Plant Biology, Cornell University, 412 Mann Library Building, Ithaca, New York 14853 USA