1. School of Biological Sciences, Flinders University, GPO Box 2100, SA 5001, Adelaide, Australia.
2. These authors contributed equally.
3. Department of Biology, York University, 4700 Keele Street, Toronto, Ontario M3J 1P3, Canada.
4. South Australian Museum, GPO Box 234, SA 5000, and School of Earth and Environmental Sciences, University of Adelaide, SA 5005, Adelaide, Australia.