1. School of Life Sciences, University of Hawai‘i, Honolulu, HI 96822, USA.
2. Department of Biology, College of Charleston, 66 George Street, Charleston, SC 29424, USA.
3. NOAA, Papahaānaumokuākea Marine National Monument, 1845 Wasp Boulevard, Building 176, Honolulu, HI 96818, USA.