1. School of Life Sciences, University of Hawai‘i, 3190 Maile Way, Honolulu, Hawai‘i 96822, USA
2. Department of Biodiversity, Conservation and Attractions, Western Australian Herbarium, Biodiversity and Conservation Science, Locked Bag 104, Bentley Delivery Centre, WA 6983, Australia
3. Department of Biology, College of Charleston, 66 George St, Charleston, South Carolina 29424, USA
4. Papahānaumokuākea Marine National Monument, NOAA, 1845 Wasp Boulevard, Building 176, Honolulu, Hawai‘i 96818, USA