1. National Institute of Polar Research, Tachikawa, Japan
2. Institute of Low Temperature Science, and Arctic Research Center, Hokkaido University, Sapporo, Japan
3. Institute of Low Temperature Science, and Arctic Research Center, and Global Station for Arctic Research, Hokkaido University, Sapporo, Japan
4. National Institute of Technology, Tomakomai College, Tomakomai, Japan
5. National Institute of Polar Research, and SOKENDAI, Graduate University for Advanced Studies, Tachikawa, Japan, and Antarctic Climate and Ecosystems Cooperative Research Centre, University of Tasmania, Hobart, Tasmania, Australia