1. Institute of Low Temperature Science, Hokkaido University, Sapporo, Hokkaido, Japan
2. Faculty of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido, and Application Laboratory, Japan Agency for Marine-Earth Science and Technology, Yokohama, Kanagawa, Japan
3. Research and Development Center for Global Change, Japan Agency for Marine-Earth Science and Technology, Yokosuka, Kanagawa, Japan