1. Utrecht Centre for Environment and Landscape Dynamics (UCEL), Department of Physical Geography, Utrecht University, Utrecht, Netherlands, and Instituto Pirenaico de Ecología—CSIC, Campus de Aula Dei, Zaragoza, Spain
2. Centre d’Etudes Spatiales de la Biosphère (CESBIO), Toulouse, France, and Instituto Pirenaico de Ecología—CSIC, Campus de Aula Dei, Zaragoza, Spain