1. Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka 819-0395, Japan
2. Department of Advanced Information Technology, Kyushu University, Fukuoka 819-0395, Japan
3. Department of Biosciences, School of Science and Technology, Kwansei-Gakuin University, Sanda, Hyogo 669-1337, Japan
4. Japan Science and Technology Agency, Saitama 332-0012, Japan
5. AMED-PRIME, Japan Agency for Medical Research and Development, Tokyo 100-0004, Japan