1. School of Marine Science and Ocean Engineering, University of New Hampshire, 24 Colovos Road, Durham, New Hampshire 03824, USA
2. Oregon State University and NOAA Pacific Marine Environmental Laboratory, Hatfield Marine Science Center, 2030 South East Marine Science Drive, Newport, Oregon 97365, USA
3. Centre for Research into Ecological and Environmental Modelling, The Observatory, Buchanan Gardens, University of St. Andrews, St. Andrews, Fife KY16 9LZ, United Kingdom