1. Department of Oceanography and Marine Geology, Faculty of Fisheries andMarine Sciences & Technology, University of Ruhuna, Matara 81000, Sri Lanka
2. National Aquatic Resources Research and Development Agency, Regional Research Center, Kapparatota, Weligama 81700, Sri Lanka
3. Department of Zoology and Environmental Management, Faculty of Science, University of Kelaniya, Kelaniya 11600, Sri Lanka
4. National Aquatic Resources Research and Development Agency, Crow Island, Mattakuliya, Colombo 01500, Sri Lanka
5. Húsavík Research Centre, University of Iceland, 640 Húsavík, Iceland