1. Botany Unit of the Finnish Museum of Natural History, P.O. Box 7, FI-00014 University of Helsinki, Finland
2. Department of Forest Ecology & Genetics, Forest Research Centre, INIA-CSIC, Carretera de la Coruña km 7.5, 28040 Madrid, Spain
3. Herbarium, Biodiversity Unit, FI-20014 University of Turku, Finland
4. Nees Institute for Plant Biodiversity, University of Bonn, Meckenheimer Allee 170, D-53115 Bonn, Germany