1. Corresponding author, Malheur National Forest, 431 Patterson Bridge Rd., P.O. Box 909, John Day, OR 97845, U.S.A.
2. Institute for Natural Resources, Oregon State University, 234 Strand Hall, 170 SW Waldo Place, Corvallis, OR 97331-8680, U.S.A.
3. Department of Botany and Plant Pathology, Oregon State University, Corvallis, OR 97333, U.S.A.
4. Colville National Forest, Tonasket Ranger District, West Winesap, Tonasket, WA 98855, U.S.A.
5. Siskiyou BioSurvey LLC, 324 Avery Street, Ashland, OR 97520, U.S.A.
6. Beck Botanical Surveys, 1708 McKenzie Avenue, Bellingham, WA 98225, U.S.A.