1. Laboratory of Plant Pathology and Entomology, Kagoshima Prefectural Institute for Agricultural Development, Kagoshima, Japan
2. Faculty of Agriculture, Kagoshima University, Kagoshima, Japan
3. Environmental Biofunction Division, National Institute for Agro-Environmental Sciences, Ibaraki, Japan
4. Agricultural Research Center for Hokkaido, National Agriculture and Food Research Organization, Hokkaido, Japan