1. Department of Cell and Developmental Biology, Graduate School of Biostudies, Kyoto University, Sakyo-ku, Kyoto 606-8502, Japan
2. PRESTO, Japan Science and Technology Agency, 4-1-8 Honcho Kawaguchi, Saitama, Japan
3. Laboratory of Subcellular Biogenesis, Institute for Virus Research, Kyoto University, Shogoin-Kawahara cho, Sakyo-ku, Kyoto 606-8507, Japan