1. Biotechnology Research Center, Toyama Prefectural University, Toyama 939-0398, Japan
2. Biotechnology Research Center, The University of Tokyo, Bunkyo-ku 113-8657, Japan
3. Division of Agriculture and Agricultural Life Science, The University of Tokyo, Bunkyo-ku 113-8657, Japan
4. Faculty of Applied Bio-science, Tokyo University of Agriculture, Setagaya-ku 156-8502, Japan