1. Division of Zoonosis Research, National Institute of Animal Health, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan
2. Laboratory of Veterinary Bacteriology, School of Veterinary Medicine, Rakuno Gakuen University, Ebetsu, Hokkaido, Japan
3. Graduate School of Veterinary Science, Osaka Metropolitan University, Izumisano, Osaka, Japan