1. Department of Genetics, Luiz de Queiroz College of Agriculture, University of São Paulo, Piracicaba, São Paulo, Brazil
2. Department of Soil Sciences, Luiz de Queiroz College of Agriculture, University of São Paulo, Piracicaba, São Paulo, Brazil
3. Laboratory of Environmental Microbiology, EMBRAPA Environment, Jaguariúna, São Paulo, Brazil
4. Department of Microbial Ecology, Centre for Ecological and Evolutionary Studies, University of Groningen, Groningen, The Netherlands