1. Laboratory of Microbial Physiology
2. Laboratory of Applied Microbiology, Research Faculty of Agriculture, Hokkaido University, Kita 9 Nishi 9, Kita-ku, Sapporo, Hokkaido 060-8589, Japan
3. Department of Biological Chemistry, Faculty of Agriculture, Yamaguchi University, 1677-1, Yoshida, Yamaguchi, Yamaguchi 753-8515, Japan