1. Department of Microbiology and Immunology, Hirosaki University Graduate School of Medicine, Hirosaki, Aomori, Japan
2. Department of Bacteriology, Hiroshima University Graduate School of Biomedical & Health Sciences, Hiroshima City, Hiroshima, Japan
3. Doctoral Course of the United Graduate School of Veterinary Sciences, Gifu University, Gifu City, Gifu, Japan
4. Department of Veterinary Medicine, Faculty of Agriculture, Iwate University, Morioka, Iwate, Japan
5. Laboratory of Zoonoses, Kitasato University School of Veterinary Medicine, Towada, Aomori, Japan