1. CAS and Shandong Province Key Laboratory of Experimental Marine Biology & Center of Deep Sea Research, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China
2. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China
3. College of Earth Science, University of Chinese Academy of Sciences, Beijing, China
4. Center of Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, China