1. Key Lab of Plant Biotechnology in Universities of Shandong Province, College of Life Science, Qingdao Agricultural University, Qingdao, China
2. CAS and Shandong Province Key Laboratory of Experimental Marine Biology & Center of Deep Sea Research, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China
3. Laoshan Laboratory, Qingdao, China
4. Center of Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, China