1. Imperial College London, Department of Medicine, London W12 ONN, United KingdombInstitute of Biology, CNRS UMR 8197, École Normale Supérieure, 46 rue d’Ulm, 75005 Paris, France
2. Imperial College London, Department of Medicine, London W12 ONN, United Kingdom
3. Hamamatsu University School of Medicine, Department of Neurophysiology, Shizuoka 431-3192, Japan