1. Department of Physics and Astronomy, University College London, London WC1E 6BT, United Kingdom
2. CoMPLEX, University College London, London WC1E 6BT, United Kingdom
3. Cell Biophysics Laboratory, Ikerbasque, Basque Foundation for Science and Unidad de Biofisica (CSIC-UPV/EHU), Bilbao, Spain